Wednesday 18 March 2020

Migratory and Non-Migratory Waders

There are generally two types of research carried out by ornithologist to collect data on birds. They are; i) field research and ii) off-field research.

i) Field research here entails the commonly used methods such as mist netting, tagging, radio telemetry and survey (example observation, interview and questionnaires).

ii) Off-field research on the other hand involves methods such as statistical analysis, phylogenetic studies and literature review (example meta-analysis).

Watching waders via field observation method can be both fascinating and challenging. It can be fascinating due to their spectacular migration pattern as well as their unique behavior adaptation. It can also be challenging due to their amazing variations in seasonal plumage and morphological distinctiveness. Birdlife International defined bird migration as the "regular seasonal movement, often North and South along a flyway between breeding and wintering grounds". These flyways are often described as 'invisible' and many theories have emerged to explain how waders find their way along those flyways i.e from the use of land markings to utilising their own built-in magnetic compass.

Waders can generally be divided into two categories i.e migratory and non-migratory waders. The word wintering and migratory are often used interchangeably and connotes the same meaning. Cambridge dictionary described migratory waders as a bird that travels from one place to another at regular times often over long distance (i.e one that regularly crosses national borders between their wintering and breeding grounds). Boreal migratory waders often breed in temperate countries and winters anywhere from the coastlines of East Asia to Sundaic Southeast Asia and as far as Australia. Non-migratory waders on the other hand are not referring to local endemics but rather migratory waders which for some reasons remain at their wintering ground throughout summer. Perhaps the description by Don Taylor in Helm Field Guides titled "Waders of Europe, Asia and North America" (2014) would best describe the meaning of a non-migratory wader:

"During their first summer, many migrant waders either remain on their Southerly wintering grounds or return only part North. This may explain the occurrence of waders summering in the Southern hemisphere that look like non-breeding adults"

Note: for comparison purposes, here is a description of a vagrant:

"In the avian world, a vagrant is defined as a bird which is often found straying into a place beyond their normal known geographical range. Vagrancy is often characterised by the bird's random and unpredictable appearance at a location. Migrants or seasonal migrants meanwhile would appear from time to time during certain seasons, and repeatedly over a period of several years"

Here is a trio of Curlews in non breeding colors.



A Great Knot with Bar-tailed Godwits



Asian Dowitcher (left) and Bar-tailed Godwit (right)



An Asian Dowitcher showing its plain underwing


Such moment will be cherished by most birders


A huge flock of Great Knots seen over Kapar coastline



An Asian Dowitcher with a bunch of Brown-Headed Gulls


"You can run but you cannot hide"

The thing about wader watching is the thrill of searching for "odd balls" among the common waders. There is a Kentish Plover hiding among the many waders in the above photo.





Most waders would appear as a whitish blob from the front.


But if you view them from the the top they can be well blended with the mudflats as can be seen from the above photo.


The current status for Curlew Sandpiper is "Near Threatened" as compared to Broad-billed Sandpiper.


Here is a Broad-billed Sandpiper among the Lesser Sand Plovers


A Lesser Sand Plover (right) and a Kentish Plover (left)


In order to get as close as possible to the waders, i would try to hide among any vegetation which sprouts out from the vast open mudflats. 


For the above photo, i hid behind a large log but there is always a comfort zone line drawn by the waders beyond which they will start moving away as shown by the photo below. 


The closest i got to them was about 10 meters (30 feet).


A Common Sandpiper seen on the mudflat. They are usually solitary as i have observed.


The underwing pattern of a Red-necked Stint appears like a shearwater.


Lots of "krabby patties" to generate enough fat and strength for their return journey.



Whimbrel


The intricate underwings of a Whimbrel


Whimbrels


Whimbrel @ Kapar


Whimbrel goes crab hunting


Eurasian Curlews


The underwings of a Eurasian Curlew looks quite plain as well.



According to Don Taylor (2014), the timing of moult is dependent on a wide range of interrelated factors such as the bird's age, the latitudes of the breeding and wintering areas, the timing of the breeding season, length of migration and the availability of food concentration. Bar-tailed Godwit  (above) and Grey Plover (Black-bellied Plover) usually molt at staging posts. Most waders that have a complete moult on the breeding grounds are sedentary species or short-distance migrants. 


Meanwhile long-distance migrant waders such as Pacific Golden Plovers (above) and Wood Sandpipers, have a partial wing moult on the breeding grounds (i.e replacing the outer primaries and inner secondaries only), and then suspending the moult until they reach their wintering areas (Don Taylor, 2014).



Here is a unique phenomenon which i have observed recently. The above arrangement appears like an unwritten hierarchy for the waders whereby the bigger waders (the curlews) occupy the lower ground area while the Bar-tailed Godwit occupies the second tier and lastly the much smaller Whimbrels at the top. Amazing !



A group of Caspian Terns with some Grey Plovers (Black Bellied Plover) and Curlews



Some of them are already displaying breeding colors.



Very soon (most likely late April or Mid May) the waders will start their journey home. Their biological instinct will tell them the exact moment to leave. 







UNTIL WE MEET AGAIN NEXT WINTER !



Sunday 23 February 2020

Lesser Sand Plover vs Greater Sand Plover

Lesser Sand Plover and Greater Sand Plover are one of those charadrius plovers which have different winter, summer and probably immature plumages to feast your eyes. Lesser Sand Plover has quite an extensive range (Birdlife International data) and over here their numbers can be six or seven times more than Greater Sand Plover. Currently there are two races of Lesser Sand Plovers. They are i) C.m atrifrons or Spot-fronted Sand Plover and ii) C.m mongolus or Black-fronted Sand Plover (Suara Enggang, Vol. 20. No. 1 (2012). The one which mostly occur here according to most field guides is C.m atrifrons or more commonly known as Tibetan Plover.

Their stunning head patterns and breast bands during summer/breeding season have kept many waders enthusiast as well as bird photographers returning to the intertidal zones. Here are some of them in their winter plumage. Photos were taken recently (Jan to Feb 2020) from the inter-tidal mudflats around Kuala Muda and the North Central Selangor coastline. Due to their winter plumage, most of the photos here appear like monochrome which makes it difficult to differentiate the various races within this species. Aside from that you can however able to see their different sizes and winter plumages from the various photos depicted here.

LESSER SAND PLOVER

A small group of Lesser Sand Plovers enjoying their early morning basking on the mudflats.


Lesser Sand Plover in flight


Their upper wings plumage can vary according to age, weather etc.




You may have noticed the different breast bands on the above Lesser Sand Plovers


From my observations, most of them would have a small patch of whitish foreheads when they first fly in from their northern breeding grounds and this whitish foreheads will spread wider while they are wintering here. More observations will be required to confirm this statement. 








They would constantly look upwards if they spot any raptors - usually its the Brahminy Kites.











Here are a few cute little ones.


Lesser Sand Plover with a Kentish Flover (third from right)


This one has an upturned 'eye-brow'.






GREATER SAND PLOVER

Here are some Greater Sand Plovers among the many Lesser Sand Plovers














LESSER AND GREATER SAND PLOVER




Here are a few charts to depict the general distribution of the Lesser Sand Plover and Greater Sand Plover in Peninsular Malaysia. These charts are by no means scientific but they do provide some baseline data on the movements of the plovers around this region. The data at Kuala Muda coastline were taken from Kampung Permatang, Keriang (5.5639, 100.3600) up to Kuala Muda coast (5.6028, 100.3489) while the info for Teluk Air Tawar-Bagan Belat (TAT-Bagan Belat) site were collated from Bagan Belat up to Kampung Batu 20. All info were extracted from entries of birders in ebird Malaysia.

CHART 1

Chart 1 shows the number of Lesser and Greater at Kuala Muda and TAT-Bagan Belat. Both Lesser and Greater Sand Plover numbers appear to be on downward trend from 2016 onwards although Lesser Sand Plover numbers show a steep upward trajectory from 2017 to 2019. The Greater Sand Plover numbers however appear to show a significant decline in 2019 (211) as compared to the numbers in 2015 (1130) i.e a substantial decreased of 81%.


CHART 2

Chart 2 shows the comparison of the number of Lesser Sand Plover at Kuala Muda Coast and TAT-Bagan Belat region. As you can see the numbers at TAT-Bagan Belat region begin to declined from year 2016 onwards and remain plateau thereafter. Meanwhile on the other hand the numbers of Lesser Sand Plovers at Kuala Muda coast has increased from year 2017 onwards. The reason for this can be attributed to the land reclamation works (to cope with the rising sea level) at TAT-Bagan Belat region which begin in late 2106 and continued through out 2017 and 2018. So it is most likely that the Lesser Sand Plover population in TAT-Bagan Belat has 'migrated' to Kuala Muda coast during that period.


CHART 3

Similarly the number of Greater Sand Plover started to plunged at both Kuala Muda and TAT-Bagan Belat regions from 2016 to 2017. Their numbers eventually increased in 2017 at Kuala Muda coast while the numbers at TAT-Bagan Belat continue to decline.


CHART 4

Chart 4 shows that while the number of Lesser Sand Plover at Kuala Muda has risen from 2017 onwards but the numbers at Mersing and TAT-Bagan Belat have declined from 2018 onwards. 


CHART 5

As expected, the coastline of Kapar has supported the most Lesser Sand Plovers in the western side of Peninsular Malaysia (37%), followed by Kuala Muda (31%) and TAT-Bagan Belat (16%). 


CHART6

For Greater Sand Plovers, Mersing and TAT-Bagan Belat area seems to share the same proportions at 15% and 12% respectively while Kapar (62%) has been the most popular roosting site for both Lesser and Greater Sand Plovers alike.

A PICTURE PAINTS A THOUSAND WORDS
A CHART SHOWS A THOUSAND MEANINGS !


HAPPY WADER WATCHING !

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